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Tilt

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Tilt

The Tilt Skull is found on the level Cortana, in the circular room where Cortana mentions that she Likes to play games too...
Its a bit difficult to get to, as it requires getting on top of the ceiling of the room. Upon entering the room, head towards the left side where you will see some mushroom shaped extrusions along the wall. Get on top of these, and then look up and towards the right for another set of extrusions. Jump to these. From here, you should be able to jump on top of the ceiling of the room, where the Tilt Skull lies in the center.